With only one remaining security shield protecting Neathia from the invading Gundalian forces, the Brawlers hatch a plan to reinstate a previously destroyed shield.
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Meicoomon rampages after witnessing Meiko Mochizuki being injured by the man bearing Gennai's likeness. Meicoomon disappears into the Real World, which has since become distorted. Possessing overwhelming power, she's the key to the world's destruction.
Upon arriving at the Digital World after the "reboot", the digidestined are hunted by a new villain. Meanwhile, Sora is troubled by her partner digimon's indifference towards her.
As Ren prepares to launch an attack on the restored Second Shield, something strange is happening deep within the bowels of the Gundalian Palace.
While Kazarina works on a secret weapon designed to take down the restored Second Shield, the Gundalians dispatch their army of hypnotized Earth kids to distract the Brawlers in battle.
On the eve of the revolt against Barodius, Nurzak sends a message to the Brawlers asking to meet and discuss a peace treaty when the Brawlers show up for the meeting Nurzak is nowhere to be found.
Fearing for her life, Zenet morphs into a doppelgänger of Ren in order to accompany Gill on a mission to destroy the Second Shield.
When Mason flees from his Gundalian ship, Nurzak and Ren are quickly in pursuit as they attempt to apprehend him before the Emperor learns of his escape.
Having made it to the generator building, Shun and Drago find themselves up against an army of Gundalian Guards.
Dan, Fabia and Shun continue to make their way through the jungle to the shield generator building, when Stoica and Jesse suddenly ambush them.
With only one remaining security shield protecting Neathia from the invading Gundalian forces, the Brawlers hatch a plan to reinstate a previously destroyed shield.
The Brawlers arrive at Neathia and meet Fabia's sister, Queen Serena with her blessing, Fabia takes her new friends to see the Sacred Orb, a source of great power that is hidden deep inside a mountain.
Speed chases Dr. Nitecol and Lorena, who has framed Speed as the culprit. Then, a fake Mach 5 appears in front of Speed. He chases the fake Mach 5 which can fly and is equipped with weapons that the original Mach 5 does not have. Speed discovers an underwater base of a gang plotting to conquer the world.
An important document is stolen from the National Science Research Center. Speed becomes a suspect as the culprit was seen driving away in the Mach 5. Moreover, Dr. Nitecol and his daughter Lorena, whom Speed was supposedly visiting during the time of the theft, say they do not know him.
The mysterious super-speed vehicle turns out to be equipped with the famous "GRX," a demonic engine that attracts racers and drives them to their death. Speed becomes fascinated by its power and takes the car, but its high-speed eventually makes him lose his consciousness.
After falling into the waterfall basin, Speed and Twinkle end up in a secret cave behind the waterfall. The two then encounter Cornpone and his underlings, who have discovered the treasure. They steal the Mach 5 from Speed and Twinkle, leaving the two in the mazy cave.
Spectra and Gus, stuck on Earth, stay in an abandoned factory. Gus returns with the new and sensational "juice in boxes."
With Ace and Marucho captured by the Vexos, Shum decides to head to Beta City alone to destroy the controller.
While Mira and Baron help out at Runo's family's cafe, Ace, Shum and Marucho head to Beta City to shut down the controller.
After realizing they have to wait to return to New Vestroia, Dan and the Battle Brawlers decide to enjoy some R & R at an amusement park while they plan their next move.
Dan, Runo, Mira, Baron, Lync, Gus and Spectra all make it through the gate and end up on Earth.
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