When the exploits of a Sailor Moon imposter become quite the sensation, the real Sailor Guardians step in to investigate. They soon learn the whole thing is a trap set by the Dark Kingdom to lure out Tuxedo Mask and seize the Rainbow Crystals.
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Sailor Venus finds the Dark Kingdom's hideout. The Sailor Guardians fall into Kunzite's trap, which hurls them into the past. The girls find themselves in a world called Silver Millennium, where they meet Queen Serenity, the mother from Usagi's past life.
Sailor Moon and the other Sailor Guardians appear to have a falling-out. But it's all part of the plan to locate the enemy's hideout. Also, a nosy journalist named Nana Asahina, who they suspect is a monster, approaches them.
Minako receives a surprise visit from Katarina, an Interpol officer from the UK and Minako's former partner while crime fighting as Sailor V. The two solved many cases together until a tragic incident forced them to go their separate ways.
The Dark Kingdom is trying again to revive the Seven Great Monsters. With five of them already under enemy control, Sailor Mercury and Jupiter scramble to protect Ryo Urawa, while Moon, Mars and Venus stay behind at Hikawa Shrine to protect Grandpa Hino.
Usagi goes on a family vacation to the hot springs, where she is unexpectedly reunited with Mamoru. But due to the Dark Kingdom's brainwashing, Mamoru no longer remembers Usagi or even who he really is.
Olympic figure-skating gold medalists Janelyn & Misha offer to coach groups of five girls. Usagi and her friends decide to investigate. Makoto, a master skater, gracefully glides across the ice and soon begins to receive personal lessons from Misha.
Usagi and her friends go to a ski resort to participate in the Moon Princess Contest, unaware that it's part of Kunzite's plan to eliminate the real Moon Princess. During the ski contest, Usagi and Rei end up falling for the trap.
Usagi decides to take the Princess Seminar course at Rose Mansion to become more princess-like, and the other Sailor Guardians join her. But unbeknownst to them, this seminar is actually a trap set up by the Dark Kingdom to find Sailor Moon.
Usagi is haunted by dreams of Tuxedo Mask, who has been abducted by the Dark Kingdom. Minako takes Usagi to a popular hair salon in the Juban District, hoping it will help take her friend's mind off of her recent worries.
Tuxedo Mask is fatally wounded while protecting Sailor Moon. Responding to Sailor Moon's tears, the seven Rainbow Crystals gather to form the Legendary Silver Crystal, and she is transformed into Serenity, the Moon Princess that everyone has been seeking.
Now that Sailor V (Sailor Venus) has joined the team, all of the Sailor Guardians are finally together! Zoisite has discovered that Tuxedo Mask is actually Mamoru Chiba and challenges him to a showdown for possession of the Rainbow Crystals.
When the exploits of a Sailor Moon imposter become quite the sensation, the real Sailor Guardians step in to investigate. They soon learn the whole thing is a trap set by the Dark Kingdom to lure out Tuxedo Mask and seize the Rainbow Crystals.
Umino seems to have fallen in love with Naru. Unfortunately, Naru only considers Umino a friend, so Usagi decides to play Cupid by getting free tickets to the amusement park and urges them to go on a date together.
Usagi and friends meet a girl who they believe holds the final Rainbow Crystal. Zoisite also pursues her. But when neither Usagi's Moon Stick nor Zoisite's Dark Crystal reacts to the girl, they realize it's her pet cat that possesses the Rainbow Crystal!
Shingo's friend Mika gives him a doll but Shingo unintentionally breaks the doll and hurts Mika's feelings after being teased by boys at his school. Shingo decides to apologize, only to learn that Mika does not wish to see anyone.
A young photographer named Kijin Shinokawa holds a modeling contest, and Usagi decides to enter. Mysteriously, all the girls who have entered the contest begin disappearing after Shinokawa photographs them!
The Juban District is having a contest to decide who can design the best handmade wedding dress. Upon learning that the prize will be a free luxury wedding reception, Usagi, Rei, and all the girls in town compete to design their own wedding dresses.
Ami brings Usagi and Rei to her favorite park where the lake and peaceful environment make it a perfect dating spot. Unfortunately, the place is soon to be demolished. Rei decides to have a date with Mamoru at the park before it closes down.
Rui is a talented tennis player at Usagi's school, but Naru has noticed a dark change in her. Worried about their classmate, Naru and Usagi go to investigate at Rui's tennis school.
Jadeite is given a final warning by Queen Beryl, who can no longer tolerate the general's continual failures. Realizing this is his last chance, Jadeite challenges the Sailor Guardians to a showdown at the airport.
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