When the exploits of a Sailor Moon imposter become quite the sensation, the real Sailor Guardians step in to investigate. They soon learn the whole thing is a trap set by the Dark Kingdom to lure out Tuxedo Mask and seize the Rainbow Crystals.
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A mysterious girl suddenly falls down from the sky, much to Usagi and Mamoru's surprise. Calling herself \"Usagi,\" the girl demands Usagi to tell her where the Silver Crystal is.
While visiting Seijuro and Natsumi's apartment, Usagi and Mamoru come under attack by the Makai Tree.
Ail and An learn that the Makai Tree is beginning to die from lack of energy. Meanwhile, Usagi and her friends visit Seijuro and Natsumi's apartment after hearing that Natsumi is not feeling well.
Usagi and Natsumi both get detention after angering Ms. Haruna during class. The two start working on their assignment when a Cardian suddenly shows up.
Usagi and the girls get into an argument over who gets to play Snow White in the upcoming play with Mamoru. After drawing straws, Natsumi gets the part, much to Usagi's frustration.
Ami and Makoto are trying to figure out who exactly the Moonlight Knight is. Suspecting that Seijuro and the Moonlight Knight may be the same person, Makoto volunteers to investigate by getting closer to Seijuro.
A school festival is going to be held at T.A. Girls' Academy, which Rei attends. When Usagi and Ami go to check up on her, they find Rei busy overseeing the setup and events.
Usagi and Mamoru find themselves looking after a baby boy while his mother is recovering from having her energy stolen by a Cardian.
Minako befriends a little girl named Mie who is a huge fan of Sailor Moon. Meanwhile, Ail and An send a Cardian to steal energy from young children.
Usagi and her friends are enjoying the cherry blossoms, which are in full bloom, when the Cardian Leshy attacks them. During the fight, the Sailor Guardians are caught, and Sailor Moon loses her power to transform.
Usagi goes to a virtual reality arcade that has just opened up in the Juban District, where she unexpectedly runs into Mamoru, as well as Seijuro and Natsumi.
Usagi, Mako, and Ami see a boy being attacked by one of Ail and An's Cardian monsters. To Mako's surprise, the boy turns out to be her childhood friend Shinozaki.
Sailor Moon finds herself facing new enemies from outer space, who call themselves Ail and An. The two aliens disguise themselves as humans to go after other human energy.
Two months have passed since the Sailor Guardians defeated the Dark Kingdom. Having lost all memory of being Sailor Moon, Usagi now lives her life as an ordinary girl.
Tuxedo Mask is fatally wounded while protecting Sailor Moon. Responding to Sailor Moon's tears, the seven Rainbow Crystals gather to form the Legendary Silver Crystal, and she is transformed into Serenity, the Moon Princess that everyone has been seeking.
Now that Sailor V (Sailor Venus) has joined the team, all of the Sailor Guardians are finally together! Zoisite has discovered that Tuxedo Mask is actually Mamoru Chiba and challenges him to a showdown for possession of the Rainbow Crystals.
When the exploits of a Sailor Moon imposter become quite the sensation, the real Sailor Guardians step in to investigate. They soon learn the whole thing is a trap set by the Dark Kingdom to lure out Tuxedo Mask and seize the Rainbow Crystals.
Umino seems to have fallen in love with Naru. Unfortunately, Naru only considers Umino a friend, so Usagi decides to play Cupid by getting free tickets to the amusement park and urges them to go on a date together.
Usagi and friends meet a girl who they believe holds the final Rainbow Crystal. Zoisite also pursues her. But when neither Usagi's Moon Stick nor Zoisite's Dark Crystal reacts to the girl, they realize it's her pet cat that possesses the Rainbow Crystal!
Grandpa Hino has become emotionally unstable, causing Rei to worry. The new apprentice Yuichiro tries to cheer her up by blaming Grandpa's behavior on the weather. Realizing that Yuichiro likes Rei, Usagi tries to help their relationship along.
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